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14-September-2008 18:02:36 - Homeostasis Homeostasis from Greek: ὅμος, hómos, equal; and ιστημι, istÄ“mi, to stand lit. to stand equally; coined by Walter Bradford Cannon is the property of either an open system or a closed system, especially a living organism, that regulates its internal environment so as to maintain a stable, constant condition. Multiple dynamic equilibrium adjustments and regulation mechanisms make homeostasis possible. The concept was created by Claude Bernard, often considered as the father of physiology, and published in 1865. Homeostatic regulation extends far beyond the control of temperature. All animals also regulate their blood glucose, as well as the concentration of their blood. Mammals regulate their blood glucose with insulin and glucagon. These hormones are released by the pancreas. If the pancreas is for any reason unable to produce enough of these two hormones diabetes results. The kidneys are used to remove excess water and ions from the blood. These are then expelled as urine. The kidneys perform a vital role in homeostatic regulation in mammals, removing excess water, salt, and urea from the blood. These are the body's main waste products. Sleep timing depends upon a balance between homeostatic sleep propensity, the need for sleep as a function of the amount of time elapsed since the last adequate sleep episode, and circadian rhythms which determine the ideal timing of a correctly structured and restorative sleep episode.1 Contents 1 Control Mechanisms 1.1 Positive Feedback Mechanisms 2 Homeostatic Imbalance 3 Varieties of homeostasis 4 Ecological homeostasis 5 Reactive homeostasis 6 Other fields 6.1 Risk homeostasis 6.2 Stress homeostasis 6.3 Waste homeostasis 6.4 Conversational homeostasis 6.5 Metabolic homeostasis 7 See also 8 References Control Mechanisms All homeostatic control mechanisms have at least three interdependent components for the variable being regulated: The receptor is the sensing component that monitors and responds to changes in the environment. When the receptor senses a stimulus, it sends information to a control center, the component that sets the range at which a variable is maintained. The control center determines an appropriate response to the stimulus. The result of that response feeds to the effector, either enhancing it with positive feedback or depressing it with negative feedback 2 Negative Feedback is involved! Positive Feedback Mechanisms Positive feedback is a mechanism by which an output is enhanced, such as protein levels. However, in order to avoid any fluctuation in the protein level, the mechanism is inhibited stochastically I, therefore when the concentration of the activated protein A is past the threshold I, the loop mechanism is activated and the concetration of A increases exponentially if dA=k A Positive feedback is a mechanism by which an output is enhanced, such as protein levels. However, in order to avoid any fluctuation in the protein level, the mechanism is inhibited stochastically I, therefore when the concentration of the activated protein A is past the threshold I, the loop mechanism is activated and the concetration of A increases exponentially if dA=k A Positive feedback mechanisms are designed to accelerate or enhance the output created by a stimulus that has already been activated. Unlike negative feedback mechanisms that initiate to maintain or regulate physiological functions within a set and narrow range, the positive feedback mechanisms are designed to push levels out of normal ranges. To achieve this purpose, a series of events initiates a cascading process that builds to increase the effect of the stimulus. This process can be beneficial but is rarely used by the body due to risks of the acceleration becoming uncontrollable. One positive feedback example event in the body is blood platelet accumulation, which, in turn, causes blood clotting in response to a break or tear in the lining of blood vessels. Another example is the release of oxytocin to intensify the contractions that take place during childbirth.2 Positive feedback can also be harmful. One particular example is when a fever causes a positive feedback within homeostasis that pushes the temperature continually higher. Body temperature can reach extremes of 45°C 113°F, at which cellular proteins denature, causing the active site in proteins to change, thus causing metabolism to stop, resulting in death. Homeostatic Imbalance Much disease results from disturbance of homeostasis, a condition known as homeostatic imbalance. As it ages, every organism will lose efficiency in its control systems. The inefficiencies gradually result in an unstable internal environment that increases the risk for illness. In addition, homeostatic imbalance is also responsible for the physical changes associated with aging. Even more serious than illness and other characteristics of aging, is death. Heart failure has been seen where nominal negative feedback mechanisms become overwhelmed, and destructive positive feedback mechanisms then take over.2 Diseases that result from a homeostatic imbalance include diabetes, dehydration, hypoglycemia, hyperglycemia, gout, and any disease caused by a toxin present in the bloodstream. All of these conditions result from the presence of an increased amount of a particular substance. In ideal circumstances, homeostatic control mechanisms should prevent this imbalance from occurring, but, in some people, the mechanisms do not work efficiently enough or the quantity of the substance exceeds the levels at which it can be managed. In these cases, medical intervention is necessary to restore the imbalance, or permanent damage to the organs may result. Varieties of homeostasis The Dynamic Energy Budget theory for metabolic organisation delineates structure and one or more reserves in an organism. Its formulation is based on three forms of homeostasis: Strong homeostasis, wherein structure and reserve do not change in composition. Since the amount of reserve and structure can vary, this allows a particular change in the composition of the whole body as explained by the Dynamic Energy Budget theory. Weak homeostasis, wherein the ratio of the amounts of reserve and structure becomes constant as long as food availability is constant, even when the organism grows. This means that the whole body composition is constant during growth in constant environments. Structural homeostasis, wherein the sub-individual structures grow in harmony with the whole individual; the relative proportions of the individuals remain constant. Ecological homeostasis Ecological homeostasis is found in a climax community of maximum permitted biodiversity, given the prevailing ecological conditions. An example of a disturbed ecosystems or sub-climax biological communities was the island of Krakatoa after its major eruption in 1883: the established stable homeostasis of the previous forest climax ecosystem was destroyed and all life eliminated from the island. In the years after the eruption, Krakatoa went through a sequence of ecological changes in which successive groups of new plant or animal species followed one another, leading to increasing biodiversity and eventually culminating in a re-established climax community. This ecological succession on Krakatoa occurred in a number of stages; a sere is defined as a stage in a sequence of events by which succession occurs. The complete chain of seres leading to a climax is called a prisere. In the case of Krakatoa, the island reached its climax community, with eight hundred different recorded species, in 1983, one hundred years after the eruption that cleared all life off the island. Evidence confirms that this number has been homeostatic for some time, with the introduction of new species rapidly leading to elimination of old ones. The evidence of Krakatoa, and other disturbed or virgin ecosystems, shows that the initial colonization by pioneer or R strategy species occurs through positive feedback reproduction strategies, wherein species are weeds, producing huge numbers of possible offspring, but investing little in the success of any one. Rapid boom and bust plague or pest cycles are observed with such species. As an ecosystem starts to approach climax, these species get replaced by more sophisticated climax species, which, through negative feedback, adapt themselves to specific environmental conditions. These species, closely controlled by carrying capacity, follow K strategies, wherein species produce fewer numbers of potential offspring, but invest more heavily in securing the reproductive success of each one to the micro-environmental conditions of its specific ecological niche. It begins with a pioneer community and ends with a climax community. This climax community occurs when the ultimate vegetation has achieved equilibrium with the local environment. Such ecosystems form nested communities or heterarchies, in which homeostasis at one level contributes to homeostatic processes at another holonic level. For example, the loss of leaves on a mature rainforest tree creates space for new growth, and contributes to the plant litter and soil humus build-up upon which such growth depends. Of equal importance, a mature rainforest tree reduces the sunlight falling on the forest floor and helps prevent invasion by other species. But trees too fall to the forest floor, and a healthy forest glade is dependent upon a constant rate of forest regrowth, produced by the fall of logs, and the recycling of forest nutrients through the respiration of termites and other insect, fungal, and bacterial decomposers. In a similar manner, such forest glades contribute ecological services, such as the regulation of microclimates or of the hydrological cycle for an ecosystem, and a number of different ecosystems act together to maintain homeostasis, perhaps of a number of river catchments within a bioregion. A diversity of bioregions, in like manner, makes up a stable homeostatic biological region or biome. In the Gaia hypothesis, James Lovelock stated that the entire mass of living matter on Earth or any planet with life functions as a vast homeostatic superorganism that actively modifies its planetary environment to produce the environmental conditions necessary for its own survival. In this view, the entire planet maintains homeostasis. Whether this sort of system is present on Earth is still open to debate. However, some relatively simple homeostatic mechanisms are generally accepted. For example, when atmospheric carbon dioxide levels rise, certain plants are able to grow better and thus act to remove more carbon dioxide from the atmosphere. When sunlight is plentiful and atmospheric temperature climbs, the phytoplankton of the ocean surface waters thrive and produce more dimethyl sulfide, DMS. The DMS molecules act as cloud condensation nuclei, which produce more clouds, and thus increase the atmospheric albedo and this feeds back to lower the temperature of the atmosphere. As scientists discover more about Gaia, vast numbers of positive and negative feedback loops are being discovered, that, together, maintain a metastable condition, sometimes within very broad range of environmental conditions. Reactive homeostasis Example of use: Reactive homeostasis is an immediate response to a homeostatic challenge such as predation. However, any homeostasis is impossible without reaction - because homeostasis is and must be a feedback phenomenon. The phrase reactive homeostasis is simply short for: reactive compensation reestablishing homeostasis, that is to say, reestablishing a point of homeostasis. - it should not be confused with a separate kind of homeostasis or a distinct phenomenon from homeostasis; it is simply the compensation or compensatory phase of homeostasis. Other fields The term has come to be used in other fields, as well. Risk homeostasis Main article: risk homeostasis An actuary may refer to risk homeostasis, where for example people that have anti-lock brakes have no better safety record than those without anti-lock brakes, because the former unconsciously compensate for the safer vehicle via less-safe driving habits. Previous to the innovation of anti-lock brakes, certain maneuvers involved minor skids, evoking fear and avoidance: now the anti-lock system moves the boundary for such feedback, and behavior patterns expand into the no-longer punitive area. It has also been suggested that ecological crises are an instance of risk homeostasis in which behavior known to be dangerous continues until dramatic consequences actually occur. Stress homeostasis Sociologists and psychologists may refer to stress homeostasis, the tendency of a population or an individual to stay at a certain level of stress, often generating artificial stresses if the natural level of stress is not enough.citation needed Jean Francois Lyotard, a postmodern theorist, has applied this term to societal 'power centers' that he describes as being 'governed by a principle of homeostasis,' for example, the scientific hierarchy, which will sometimes ignore a radical new discovery for years because it destabilizes previously-accepted norms. See The Postmodern Condition: A Report on Knowledge by J.F. Lyotard Waste homeostasis Andrew Potter has used the term waste homeostasis in reference to the lack of net gain from energy-saving technologies.3 Conversational homeostasis A 2007 study purported to find and show clinically conversational homeostasis in which overly-familiar people such as spouses condense their speech so much that they are actually worse at communicating novel information than strangers are, while not being conscious of this problem. 4 Metabolic homeostasis Some herbal medicines, known as adaptogens, have been defined to function as non-toxic metabolic regulators that can enhance metabolic homeostasis during stress.5 See also Acclimatization Allostasis Apoptosis Biological rhythm Claude Bernard Cybernetics Enantiostasis Gaia hypothesis Health Homeorhesis Metabolism Osmosis Protobiont Self-organization Aging Steady state Lenz's law Le Chatelier's principle References ^ Wyatt, James K.; Ritz-De Cecco, Angela; Czeisler, Charles A.; Dijk, Derk-Jan October 1999. Circadian temperature and melatonin rhythms, sleep, and neurobehavioral function in humans living on a 20-h day. Am J Physiol 277 4: R1152-R1163. Fulltext. PMID 10516257. Retrieved on 2007-11-25. ... significant homeostatic and circadian modulation of sleep structure, with the highest sleep efficiency occurring in sleep episodes bracketing the melatonin maximum and core body temperature minimum ^ a b c Marieb, Elaine N. Hoehn, Katja 2007. Human Anatomy Physiology Seventh ed.. San Francisco, CA: Pearson Benjamin Cummings. ^ Potter, Andrew 2007, Planet-friendly design? Bah, humbug., MacLean's 120 5: 14, http://www.macleans.ca ^ Keysar, Boaz 2007, The Effect of Information Overlap on Communication Effectiveness, Cognitive Science, http://www.eurekalert.org/pub_releases/2007-02/uoc-wec022007.php ^ Winston, David Maimes, Steven. ADAPTOGENS: Herbs for Strength, Stamina, and Stress Relief, Healing Arts Press, 2007. v d e Subfields of and scientists involved in cybernetics Subfields Polycontexturality · Second-order cybernetics · Catastrophe theory · Connectionism · Control theory · Decision theory · Information theory · Semiotics · Synergetics · Biological cybernetics · Biosemiotics · Biomedical cybernetics · Biorobotics · Computational neuroscience · Homeostasis · Management cybernetics · Medical cybernetics · New Cybernetics · Neuro cybernetics · Sociocybernetics · Emergence · Artificial intelligence Cyberneticists Igor Aleksander · William Ross Ashby · Anthony Stafford Beer · Claude Bernard · Ludwig von Bertalanffy · Valentin Braitenberg · Gordon S. Brown · Walter Bradford Cannon · Heinz von Foerster · Charles François · Jay Wright Forrester · Buckminster Fuller · Ernst von Glasersfeld · Francis Heylighen · Erich von Holst · Stuart Kauffman · Sergei P. Kurdyumov · Niklas Luhmann · Warren McCulloch · Humberto Maturana · Talcott Parsons · Gordon Pask · Walter Pitts · Alfred Radcliffe-Brown · Robert Trappl · Valentin Turchin · Jakob von Uexküll · Francisco Varela · Frederic Vester · Charles Geoffrey Vickers · Stuart Umpleby · John N. Warfield · Kevin Warwick · Norbert Wiener Retrieved from http://en..org/wiki/Homeostasis Categories: Biology | Cybernetics | Physiology | Systems theoryHidden categories: All articles with statements | Articles with statements since February 2007 Views Article Discussion this page History Personal tools Log in / create account Navigation Main page Contents Featured content Current events Random article Search Go Search Interaction Community portal Recent changes Contact Donate to Help Toolbox What links here Related changes Upload file Special pages Printable version Permanent link Cite this page Languages اردو Bosanski БългарÑ?ки Català Česky Dansk Deutsch Eesti Español Français Galego í•œêµì–´ Õ€Õ¡ÕµÕ¥Ö€Õ¥Õ¶ Hrvatski Ido Bahasa Indonesia Ã?slenska Italiano עברית Lietuvių Magyar Bahasa Melayu Nederlands 日本語 Plattdüütsch Polski Português Română РуÑ?Ñ?кий Simple English SlovenÄ?ina SlovenÅ¡Ä?ina Suomi Svenska ไทย Tiếng Việt УкраїнÑ?ька ä¸æ–‡ This page was last modified on 13 September 2008, at 14:06
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